The enigmatic Hyloxalus Edwardsi species group (Anura: Dendrobatidae): phylogenetic position, a new species, and new putative morphological synapomorphies - Anganoy-Criollo, Marvin; Viuche-Lozano, Andres; Enciso-Calle, Maria Paula; Bernal, Manuel Hernando; Grant, Taran (Herpetologica, 2022)